| 1 | LE, FUT3 |
| Five novel missense mutations of the Lewis gene (FUT3) in African (Xhosa) and Caucasian populations in South Africa. | |
| Pang H, et al. | |
| Hum Genet 102 : 675-680. 1998 | |
| 2 | C3, FUT2, LE, PEPD, BCAM |
| Genetics and linkage relationships of the C3 polymorphism: discovery of C3-Se linkage and assignment of LES-C3-DM-Se-PEPD-Lu synteny to chromosome 19. | |
| Eiberg H, et al. | |
| Clin Genet 24 : 159-170. 1983 | |
| 3 | LE |
| Probable genetic linkage between the loci for the Lewis blood group and complement C3. | |
| Weitkamp LR, et al. | |
| Cytogenet Cell Genet 13 : 183-184. 1974 | |