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GrailEXP v3.31 [March, 2002]                   http://compbio.ornl.gov/grailexp/

Authors:  Doug Hyatt, Manesh Shah, Victor Olman, Richard Mural, Ying Xu, and 
  Edward C. Uberbacher, 1996-2001

Reference:  "Automated Gene Identification in Large-Scale Genomic Sequences",
  Xu, Y. and Uberbacher, E.C., Journal of Computational Biology, Volume 4,
  Number 3, 1997

Sequence:  >GrailEXP Input Sequence (106643 bp)
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PERCEVAL Exon Candidates (26 predicted)

 Index Std   Begin       End     Frm    Type     Len    Scr    Quality

Exon  1 +        412        581   1   Internal    170    59    Marginal
ttctcaaatgtccctgttcctgaattactcagagggctaatccccagtttctctgtgcct
ccacagctgcctgtcatttcatttatcccaaagtggcattcccatttgtttacctgtcta
ctaggatactcctagaatgcaggtagcatgagggaagagccttcactcatctttgtgtag
ACGGCATTGTGACCCTGGTGACCAACCCTGGCACACATCAGAATCATCTGGGAGCTTTTC
AAAAATATGAATGCCAGAGCTCACCCCTGAACCTACAGGAACTGAGCCTTACACATCCAT
ATTTTAAAATGTCCCCACATCACCGTTGCACACTGTTCTGCACAGAGCAG
gtatttttaatagttctcgaatgtctagtcaattacttagcaaggctgcctggagaatca
tgtctaccttagcttacttctccggctttgtaggtttttgttttcttttgatttcagttt
ccatttcctctaacttgaaagtatgtgcattggtaacagccacacagcagtcattactga

Exon  2 +       1266       1453   0    Initial    188    57    Marginal
ggagaatacgtggggaagttctctggggcgggcgcagtctgctgctgattctgagaaacc
gaaactcagctgttaaaaccgctgcctctgcactttggaatcccatcttgagactcgcta
agcgtcccagccgcatccctcccgcagcgacggcggcccgggacccgcgggctgtgaacc
ATGAACACCCGCAATAGAGTGGTGAACTCCGGGCTCGGCGCCTCCCCTGCCTCCCGCCCG
ACCCGGGATCCCCAGGACCCTTCTGGGCGGCAAGGGGAGCTGAGCCCCGTGGAAGACCAG
AGAGAGGGTTTGGAGGCAGCCCCTAAGGGCCCTTCGCGGGAGAGCGTCGTGCACGCGGGC
CAGAGGCG
gtgagtattttaacaacctggcttcggggcgggagcccgggccccaaacatactcactca
caagcaccttggtccttggagtttaaaaccacaggggcccaacaacagggattttgactg
aatgaaagctcatttgcttaaatgaaagtatccatgccgcaccttataaattgagtcggc

Exon  3 +       7177       7297   0    Initial    121    48    Marginal
gcatagctggtttttctagctaccagtctcttaagctagctgtctagggacctcaccaac
agacattagcataaaaaagacacttagcactctgggaatcccaagggtttggggagctct
gtcccaggaaccagggaccgtggctaaatatttattttttattatgtcacaggagtttaa
ATGGCCTCTGGACAGCCAGAGAAACTTGTTATTCAGATTGTCTTCAGAAGAACTTACAGT
GGGGAGCATAGTCAGCTGACGGCCCCAGCTGCCACCCTTTTGGATCCATTACACCTTGCC
T
gtgagtgctggcttgtgctgcacccatcccagccaatggctgagcattccagggtgccag
tgctgttgcaggactgccaggcttatacgccctctgggcagcaacagacccatacactga
caatggcacttgcagcagagaaagagtttaatcatcgcgacattctcagtgaggaaacag

Exon  4 +      10845      10958   1   Internal    114    50    Marginal
gagaaggtcagagaggctttcctccttttgctgttttctcaaataccaaggtgctatatt
ttgggataacatgtcgtgaaccccatcatagctttaaaatgatttgaatagagaagtact
ttgcaggagaaaagtttctctggggattccctttgttctgctgcttttcacaaatgacag
GTGAGCTTGGTAGCCTTGTGTTAGTGTTACTGGCAAGGGGTCCTGATCCGGACCCCAAGA
GAAGGTTCTTGGATGTTACACAAGAAATAATTCAGGGCAAGTCTATAGAGCAAA
gtgagagcaagtttattaagaaagtaaaggaataaagagtgactactccataggcagagc
agccccaaggcttgctggttgcccatttttatggttatttcttaattttatgctaaacaa
ggggtgggttattcatgcttcccctctttagaccatctagggtaacttcctgatgttgcc

Exon  5 +      13350      13486   1   Terminal    137    64    Marginal
tgattgcactccatcatatagacggatcttgtttttcactttggcttattatttagttgc
acaaggaaacaacctttgttctcacagtggcatcttggataacatgtcttatctagtatt
ctttctaaaaatgtttaacctaatcgaatcatgaggaaataatcttttccttacatccag
ATTGTGAAGCATTCTATAAGAAAAGTTACCTGAACTCTTCAAATATGTCAGTCATGAAAG
AAAGAAAAAAAAAGTCAAGGAAATTGTCTGAGCTCATAGGAGTCTACAAACACATGTCAA
ACCATAGAAATGTGTAA
tccataattggatctaggatgaggtaaaggtagggaagaatattaaaatggaaacaattt
ggaaatttgaatatagactattaggcaatattaatgttaaatttcttggtgtgataatgg
tattatatttaagtaggagactgtttttatttttaggagatacatgctaaaatatttagg

Exon  6 +      21054      21157   0   Internal    104    69    Marginal
tcaaaagtcttagactggagtctcatcagtgttgctcaattgacctggtgatctttggcc
tgagattcaacctctgtgtctttaccagtgatttatgaatacctaagttttcttttatcc
caaatgttctaagatagcttgattgctctatagtagtggttttccaagtgtggtacctag
ATCAAGAACAGCATCATCTTGGAACTTGTTAAAAATGTAAGTTATTTGACCCAGTCGCAG
GCTACTGAATTAGAAACTCTCAAGCTCGCGGACAAGCCCTCCTG
gtgattatgatgcacgataaaggctgagaactactgatctggagtatagtcaaatttgtg
gtatagagagaagctctataccacaatgttagcagacattgaagattgtgtagcaatttc
atgattggcatttacgtctcgtatctgatgtttcagctttcagtctctgtagcatggctc

Exon  7 +      22258      22337   0   Internal     80    52    Marginal
aggggcttggaatacacatcaatcaagcacactctctgccctcaaaagctcacaatattt
taacagcttcctagtgtcacagaggtgaatatttttcttaaaaagtcttccctgatccgc
ctggttgaaatgactccttgcttctccgagcccctgattatgctcaacctgtgtccctag
GAGAAGAAGAGACCAGTGGACATAGGCCTAGAAAAGTCAAAGCCTAAAAGCCAGTTAGTT
ACTCAGATCATCAAGAGGAG
gtggctgcattgcaaatggcctctaggatcagtctgtgggaacagaaaatgtaccaatgt
ttcctgtaccccggtgtggtcacacagaagagaatgccaatctaaagccatcttaaagga
atttactcacaagcaccacctagaggaacggtggagccctacaaggacagtacaccaact

Exon  8 +      23221      23415   0   Terminal    195    48    Marginal
agatcctggatttctgttggcttgaacaactttacacttattgccaataaacctccttga
gggtagggaccacatcttactcgtttttgtgctctctgtacttaatgcgtacataacatg
tttaattgatgtacagtaaatataaatgctgccatttaaagttatagcattatcctttag
AGGTGGAAGAGAACTAAAGTTCATTTAGTTCACAAAGAAATTGTAGGCTTGCTCAAGTCC
AGCTGTGGGGCTATGCTGAAATTCGATGGTCTCCTTTATTACATGATTGGTAGGAAATTT
AACTTCATGACTAATGATGGACAGAAAGGCACAAAAGGGAAAAAGCTCTCTAAAGAGTCT
CCACCGGACATGTAG
ctccgggtgggacagcgggtcagtgcagaatatgacaatcacagccatgcagggactatg
ggaactgccagcagttctggtatcttacaagcttttgatttaaatacttaataggagatt
aagtctcaatgtggtctggatttcgaaatgtttcagaatttgtttttctagtaatggaca

Exon  9 +      24254      24337   1   Internal     84    94   Excellent
gggcccaggatgacctgtacgggtcagggctgaatattgtgtttctaacctggaagcctt
gttcactcatgctcacatatagttaattgccttctaatagttgtttcttatgctttgaag
ggactggttttcttttagaagtctaataattgacgatttgatatttctcgcctggcttag
TTGCGGAGCAGGAGCTGGCCAACCTGGAGAAGTGGAAGGAGCAGAACAGAGCTAAACCGG
TTCACCTGGTGCCCAGACGGCTAG
gtaagcaagcacagccgctgccggaatgagggagggcggggccacctatggatcctgtaa
gcagcttgcatatgtctgtctgtgaatgtccattgtagtttgtttggtcattgtgctgct
tgggtggaatgcatttaattgtgtgtgtgtgtgtgtgtgtgtgtgtgtgtgtattttttt

Exon 10 +      25666      25780   0   Internal    115    81        Good
gctctaacccaggcctgtttcaggtaggaatccagcaagtagaacagggtgtggctggac
ttttctctattcatggctctgttcccagattctgacctttctgggattgaagggggaagt
ggagggagataagagagttcggggttcgtaatgcagtgatccttgtttctttgagcctag
GTTTACACTGACTGCTCGTTCTTCTTCCTGGGAATCTTCGAGTTTTTCAGCATCCCCCAC
TCTCTGCTACCCATCTCCCACCTGGATTTCCCTGATACAGACGATTTCCTCTCCT
gtcggcagggcttggcctcactgttggctcttggtagactccttttagggttccattgtt
cttgacatggccctcataggcaagtccctttagatggctttaggttggcattctctttca
tgagaatcacattaaaagtacaggaaatacttgtgcattctttcttttcataaactgtga

Exon 11 +      30093      30176   0   Internal     84    79        Good
gcagaagacgtagcccggccattctctgtgatctaagtagcttgaagtaacaacctataa
tcatggtgaggtatagcacaaacaatactgtaatttgcctttcatcataattttgtgttc
cagagttcagactgaaaaatggtaactgtttttgttctctgtaatttcaatacaatttag
CTAAAAAGAGAAGAATCTGTAAGAATCAAGAAGGAAGCTGAAGAAGCTGAACTCCAAAAA
ATGAAGGCAATTCAGAGAGAGAAG
gtaaggaggaggaggattagttcacatttgctgatcaacaattatagtttcgttgtagaa
gatttttattatagtaactttaaaaatttatccatctttctctctctctctctttctatt
ccccactcttcttttcctcatccagtttttttccttccattttatccttcaaaggaagat

Exon 12 +      34051      34143   1   Internal     93    76        Good
agagctgcatacaggtacatgttaagggatcacacttgttgctttcttatttctaggaag
caagtagtagcaagggctaacagtttaagaacaattttaggggtaaactcaacattgagg
aagttatttcaatgttgtttatataaattgggaaacatttccatttctgaaaactttcag
GGAACATCAGTGTTTCCAAGAAATGTGGCTCCTACATCATGCATTTCTTTGTGGATGACT
TGGTGGAAGCATTAAGCCTTTGGTTATTGAAAT
gtaataggatctcgctaattaaccccataatgtggtaatgttagtatctctaaaataaac
atgtggtactaagagtgtaggctgatagcttttctgtgcttcacttaccttatttgtaaa
atgggtataataatgtaatagtgagactggtaggactagaggatgcaatatatacacagc

Exon 13 +      36718      36813   0   Terminal     96    71        Good
gcacatacattcctatataattattccttcttactttctcatccttatcttctactcctt
tctctataaacgttgttcaatttatccatgagcttcttgggtttccttgagctaacatat
tctcttgtacctttgtatattaaaaacgttaagttctgggaatgcccctttcctgtccag
GGTTACTACCAGCCCATGGTTGCCTTTGTGCAAATTAGAGACCAATTCTCCATCCAGGCG
GTCAATTCTGTGGGTGCTCAGACTGGTGATAGATAG
tgcctttgtgagacacagaaaaagacacctcctctgcgctaaagcagccacaagctgcag
tacaaatcttggtgagctgaacactggctctatttttcctttcctactcccctcctgcct
gctgggaccccatgggctgaacaacccatataatggagtgtagtaaccctgttcctacct

Exon 14 +      40962      41100   0    Initial    139    44    Marginal
cttagcctgtctgtcccaactcttcttctaaagaaggcatttgagtgaggagctggtggg
aagctaaggagctaagtaccaggagaaactgtctgtcttgcagagagatatggtgggagg
gaggcaccagatgaagactgtgaatgtatcattgcctctcctttcatgggtagagaatga
ATGAGAATTTATTTTGAGGCTCAGCCGGAAGAGTTGAGAGAGATTAGGGGCTGGGCTGGT
GGCTTTGTGTCTGGAGTTGGTTCCTTCTGGTGGGCTCTTGGACTCGCTCACTTCAAGAAT
GAAGCCATGGACCTTTGCG
gtgagtgttacagctcttaaaggtggcacagacccaaagagtgagcagcagcaagattta
ttgtggagagcgaaagaacaaagctttcacagcatggaaggggacccaaacaggttgccg
ctgctggctggggtggccagcttttattgccttatttgtctgcgcccacatcccgttgat

Exon 15 +      50200      50274   0    Initial     75    57    Marginal
tagtgtttggaaagagacgctgaatacagtgtaactccaacaaggacagcagaggaagaa
gtgaaggaaagtgagcagccaactccagcctttacccccaacaatagagcctgagaactc
ttcttgatgcataagattcactaagaaggccctggggcccagaaagtgcttctggtaaac
ATGCTATACCTGGGGGAATATGTGGTCCCCATTCTGGAGACAAGCCATCTGCATAGCCAA
CTTCATCACTTCTAT
gtgagctctaagagccgtattttaaaaattacatgaaaatgaatgctagtcaaacaagta
gtaaatccaagaaagaggaaggcatagataccagaacaattgtgaacaaagaagacagtg
acatttttggtcaaatatgtgcttgtttttctgtttaaataagaatcagggctgggcgcg

Exon 16 +      50313      50391   0    Initial     79    46    Marginal
agaactcttcttgatgcataagattcactaagaaggccctggggcccagaaagtgcttct
ggtaaacatgctatacctgggggaatatgtggtccccattctggagacaagccatctgca
tagccaacttcatcacttctatgtgagctctaagagccgtattttaaaaattacatgaaa
ATGAATGCTAGTCAAACAAGTAGTAAATCCAAGAAAGAGGAAGGCATAGATACCAGAACA
ATTGTGAACAAAGAAGACA
gtgacatttttggtcaaatatgtgcttgtttttctgtttaaataagaatcagggctgggc
gcggtggctcacgcctgtaatcccagcactttgggaggccgaggtgggtggatcatgagg
tcaggagatcgagaccatcctggctaacaaggtgaaaccccgtctctactaaaaatacaa

Exon 17 +      54300      54477   0    Initial    178    82        Good
aagttctcaaaggaaattaaaaatgctactccagtgaacatgtgaatgatacgaaggcaa
aatagctgtattgctaatatgaagaaagttcaagtggtctggatagaagatcagaccagc
caaaacattcccttaagccaaagcctaagctagagcaaggccctaacactgtttaattct
ATGAAAGCTGAGAGAGATGAGGAAGCTGTAGAAGAAAAGTTAGAAACTAGCAGAGGTTGG
TTCATGAGGTTTAAGGGAAGAAGCCAGCTCCATAATATAAACATGCAATGTGAAGTAGAA
AGTGCTGATGTAGAAACTGCAAGATATCCAGAAGATCTAGTTAAGATCATTGATGAAG
gtgtgtatactaaacaacagattttttttttttttttttgagacagagtcttgctctgtc
tcccaggctggagtgcagtggcgcaatctcagctcactgcaacctcctcctcccaggttc
gagtgattctcctgcctcagcctcctgagtagctgggattacaggcacccaccaccatgc

Exon 18 +      55583      55707   1   Internal    125    93   Excellent
ttaatgttttcatgcttgctaacgcagtctccattctgcagcctatggatcaaggaataa
atctgatgttcacatcttattatttaagaaatagatttttaaaggctatagatggtgatt
cttctgattgatctgggcaaactaaattaaaaaccttctagaagggatttaccattctag
ATGCCATCAAGAACACTTGTGATTCATGGGAGGAGGTCAAAATATCAACATTAACAGGAA
TATGGAAGAGGATGATCCCAGCCCTCATGGATGACATTGATGGACTCAAGACTTCAGTGG
AGGAA
gtcttgcagatgtggaaatagcaagagaactagaattagaagtggagcctgaagatgtga
ctgaattgttgcaatcttatgatcaaacttcaacggatgaggagttgctttttatggatg
agcaaagaaagggatttcttaagatgggaactactcctgctgaagttgctgtgaacattg

Exon 19 +      56375      56542   0   Terminal    168    94   Excellent
ttagcattttttttagcaacaacgtgtatttttaaattaaagtgtacatttttcaagaca
taatgctattgcacacctaagagactacaatctagtgtaaacccaacttttatatgaatg
gaaaacaaaaaaatttatgtgactcactttattgcaatatttcatttgctttttttgcag
TGGTCTGGAACCAAACTCACAATATCTCTGACGTATGACTATAACTACATATTGTATGTA
AGTTCAATAGGACATATTGATATTGGGGATGAAGAAGGAGGAAGAATCAAGGATGACTCC
CAAGTTTCTGACTTGAAGTACTGGAGGTGTAGTGATGCCATGTACTGA
gatgggggctactaggtgaaaaccaaggtttgggaagaaaaatatcaagagtttaagtta
agtttaagtttacatatttagaaaaatatcaagagtttaagtttgagatggtttggatat
atcaaatgggcagttgtacacacgaatttggtctcactgagcagatatgtgggaaatata

Exon 20 +      56903      57023   0    Initial    121    45    Marginal
aatttaggaattatattagcatgtaaatataaagccataaaagtggacaaagtcacttaa
gatgaaattgcaggtttgagaagagggcccaggaccaaatacagaggagctctatcatgt
agagttcgggtagatacgaaaggacatccgagaagtcgggcaaattaagagtgtggtatc
ATGGAAAACAGGACAGACATATTTCCGAAAGGAGAAATTAGTCAATTCCTTACTAGTGTT
GAAAGTTCAAATGAGAGAAGGACCAGTGACAGTGGCTGTGATCTTGACATAAATGATTTG
G
gtatgatgatggggctggagcttgattgaattggacttacgagtaaatggagatatggga
atagatttggcatttatcagtaactcttgaataatttgaaaatgagaacaatgaaatgag
gcagtaactggatgaggctgcggaatcaaaagttttcttaaattgatcatgtttgtatgc

Exon 21 +      58961      58998   0    Initial     38    44    Marginal
ggaccattacaacgtccccccaaccacaaccccagcatgaagacaattgttagaaaggtc
aggaacaggtactaaataagaaaaatcaagtctataaactggaagtgtccaaggtcatca
gtgtaagtctagtacaaatccaacaataccaccaaatcgaccagaaacgcagtgctcaca
ATGGCCATGGCAGCAGCCAAAGCCGTCAACATCAAGAG
gtagatgagaaagtattaataactggcgttccaagcccagatgcagttgttcacccaaac
ctaacgatggtcaaaacggtgcacacactgcagtcagtgctttgatcgagctggtgttcg
taactcataagtagagcacctcacctctttagaaacatcactgaattcataaacttgaag

Exon 22 +      60554      60644   2   Internal     91    56    Marginal
ttatgattccttaatttcctccaatatgtgacagtacctcattttttcccttctctttca
tgaccttaacacattcagtgggtaatggatagttactttgtagacttcccccagtttggg
tttgtttgatgttttctcatgaatagcttgaagttataatttccaacaagaaaattacag
AAGTGATGCCATGTCCTTCTCAGTGCATCGCATCAGAGGGTACATGATGATCAGTATCCT
TATCACTGGTGATATTTACTTGGATCATTTG
gtaaggtggtctccactgttaagttattgtttttctctttataattgatgaatgttttgg
gggagatattttaagactatactaatatcctgttatctgcaaacttttgcccagtgattt
tagcctatgtaaattaatcttgccaacatttttttttttttttacctggatgtttgccta

Exon 23 +      67516      67605   0    Initial     90    44    Marginal
cacccacccatccacccatccattcttccatccttccatccttcctaccttccttccttc
cttccttccatccatgcatccatccatccttccttccattctttaaattcataggtctat
ataaggcctactaagtgcctacagtttctgcccaatgaaccttagtaaaggagaacacta
ATGGTTTTAAAGCAAACTGCTGAGCACAGAGCATATCACAATAAGAAAGCACTACTGCCT
TATCAGGAAGAAGCATCTAAGTCAGACCAT
gtgcgttagatgcttcttgggaaatgacaccagagcagaagcttaaactatgaatcagta
aagtgatgagtctttccaggtatcagaattaataaagacacatagagcaagaattaaggt
ataaatgcaagaaataagcatggcttgtgttgcggaagaagagttatgtagctgcagtgc

Exon 24 +      75053      75115   0   Terminal     63    67    Marginal
tgagaaaagttacataaaagttcacgtaggctggaaaaagggggaaccagcctacctgaa
ctttatggattattagtcctgagaactaggtctgaaatatgtaacctgctgccatgaagc
tgctgatagttttttatgagatccatgtaatttcctaagcattagctctttgccttacag
AAGCTGGGGCTGCAAATCAATGTGAAGGAGAGAAAGCATATGACAAATAAACAGAACACT
TGA
aatgccataaatgtcagaacacatggtcactatgccagggtgtggaactctggtgtggtg
ggtcagagacaccgaagggcattctccctcatgtgctggcttgccaaaggctttgttgtc
atgtgttcctgaataggctttgttttccttttcagaggtagaaataaaacttcatccaga

Exon 25 +      80891      80935   0    Initial     45    54    Marginal
ccaagattgctgatttggtaagtagcagtgttctgctttgtaagtgcaatattctttctc
atctcatcttatttaatgtttgtatatttgtaactatacaaacataagttttctatttac
catattaactgtttttttctttccattttaatctttatctcccatggaattctttctcaa
ATGTTTGATTATTCTTATATGTATTTGGATATGCTGCTGAATGAG
gtatgtaggagcttgtcttctgtgtggagtagactcgtgttctcttatagtcagtaggta
cttgggacgttgccctgtttccctgagcccagccatgcctctctcattcacttcttacct
tatgggccatgatagtgctgcctactatgtagtgcaagtgtgggagggaaaagaacccac

Exon 26 +      93048      93125   0   Terminal     78    92   Excellent
ccacagttatttgctacctacgtatataagaaaataatttcacatccaaatagtacttaa
gtaaaagtttcaaaaatgaaatattgtaccagttgaggtttaaatatgtaattgatttgg
gtatcttctgtacaactgtgttaattgaaaaccatattttaaaacttttattgttcctag
AGTGAATTACTGGAACTGAAACGGCAGCAGCAAGAGCAAGAAAGAGCCAAAATCCACCAG
ACTGAACACAGGAGGTAA
aactggctaataaaataagcatccttggttaaatcaattacctagaaagtttgtacagtg
ttttaggaaagcattacgataacattattaatatggttttggaatgcgtgttacctttaa
atctgttattgggacgtgatatatgtagaggtatgccaactgctacatatataagtacac

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